Vd. Sensory Systems: Chemical Senses



OLFACTION


STIMULUS
Olfaction is the response to chemicals that contact the membranes inside the nose. Olfaction is important in food selection, as most of what we call 'flavor" is a combination of the taste, texture and aroma of the food. The stimulus for olfaction are air-borne molecules that are ordorant chemicals-

TRANSDUCTION
Olfactory cells line the nasal cavity. Each ell has cilia that extend from the cell body into the nasal passage. Olfactory receptors are located on the cilia.  Ordorant chemicals activate synaptic transmission. Olfactory receptors have synapses with cells in the olfactory bulb.

While there are only 3 types of cone cells that produce color vision and 5 types of taste receptors that provide flavor, there are several hundred types of olfactory receptors that respond to different types of chemicals. Olfactory receptors are vulnerable to damage since they are exposed to the air. The lifetime for such cells is just around a month.



TRANSMISSION
 When an olfactory receptor is stimulated the axon synapses in the olfactory bulb. Coding for the smell begins in the olfactory bulb. 

Cells of the olfactory bulb axons synapse in the olfactory area of the cerebral cortex known as the olfactory cortex.






SENSATION PERCEPTION COGNITION
Olfaction also plays a role in social behavior.  What is the name of the scent-related chemicals involved in social behavior?  Who detects orders more readily? Men or Women 

SYNESTHESIA
     What is synesthesia?  What sensory systems are identified in the textbook explaination of synesthesia?  


 PUT IT ALL TOGETHER
For the final exam, you should know the answer to the questions in red above.  Also you should be able to associate the following terms within the correct cell of the class schema.

air borne chemical ordorants
medial dorsal nucleus of the thalamus
nasal passages 
Olfactory bulbs
olfacotry glomeruli
Olfactory protein molecule
olfactory receptor cells
orbitofrontal cortex
a rose

GUSTATION

CHEMICAL CODING
Most evolutionary scientists believe that the first sensory system of the earliest animal was a chemical sensitivity.  Before the neocortex (Forebrain) evolved, there were areas of the midbrain and hindbrain devoted solely to sensing.  The cranial nerves are illustrated on pg 85 (unit 5.7) of the coloring book.


STIMULUS
 Organic substances containing chemicals that are perceived as sweet-sucrose, sour-HCL, bitter-quinine, salty-NaCl and umami or meaty-MSG.  A combination of activity in five kinds of receptors (along with smell) give the perception of the taste of the food.

Taste buds are contained in papillae Taste receptors have excitable membranes and release neurotransmitters to excite other cells. Taste receptors are regenerated and replaced every 10-14 days, which is why food may taste more intense after an illness or after an extended fast.  Taste receptors are located inside of tastebuds which are located in the papillae of the tongue.

TRANSDUCTION
When a substance tastes salty for instance, it means that a saltiness receptor has detected the presence of sodium. Sodium ions cross into the membrane and produces an action potential.  Sweetness, bitterness and umami receptors resemble the action of metabotropic receptors and therefore activate G-protein molecules and second messengers within the cell.







TRANSMISSION
Although each receptor detects one kind of taste, several receptors create a particular firing pattern that is perceived as a single taste experience. Information from the receptors in the anterior 2/3 of the tongue synpases with the Chorda Tympani nerve and information from the posterior tongue and the throat travel along branches of the Cranial Nervs IX and X.  What are the names of Cranial Nerves IX and X?











 Taste nerves project to the nucleus of the solitary track (NTS) in the medulla. From the NTS, the information branches out to the pons, the lateral thalamus, the amygdala and the ventral posterior thalamus (VPN), finally terminating into areas of the cerebral cortex. The somatosensory cortex is where touch or food texture is detected on the tongue and the insula or primary taste cortex is where taste is perceived..  Innervation is ipsilateral in this sensory modality. 








SENSATION, PERCEPTION AND COGNITION
  The sensation of taste can be affected by culture and familiarity but also by genes and hormones. Give an example of how genetic differences affect taste. Also how do hormones affect taste preferences.



 PUT IT ALL TOGETHER
 You should be able to associate the following terms within the correct cell of the class schema: (Stimulus, Transduction, Transmission and Sensation Perception and Cognition).

 NaCl
Nucleus of the Solitary Tract
papillae 
Primary gustatory cortex
saliva
taste buds
taste receptors
teeth
tongue 
unami
Vagus Nerve
Ventral Posterior nucleus

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